{"id":25820,"date":"2021-03-03T15:30:59","date_gmt":"2021-03-03T15:30:59","guid":{"rendered":"https:\/\/www.ispa.pt\/oferta-formativa\/doutoramento-biologia-do-comportamento\/"},"modified":"2025-11-03T16:27:57","modified_gmt":"2025-11-03T16:27:57","slug":"doctors-behavioural-biology","status":"publish","type":"oferta-formativa","link":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/","title":{"rendered":"Behavioural Biology"},"content":{"rendered":"\n<p class=\"has-medium-font-size\"><strong>School of Biosciences<\/strong><\/p>\n\n\n\n<p class=\"has-medium-font-size\">This programme aims to provide advanced training in the theories and methods for studying and analysing the main research areas in Behavioural Biology in which Ispa has recognised expertise and excellence, namely: neuroendocrinology and the ontogenetic development of behaviour; behavioural ecology and conservation; behaviour as a factor in evolution; and animal behaviour and welfare.<\/p>\n\n\n\n<!-- EXPAND -->\n\n\t\n\t\t<a href=\"#\" class=\"expand-header js-expand\">\n\t\t\t<div class=\"expand-header-text\">\n\t\t\t\t<h2 class=\"expand-block--title\">About the Programme<\/h2>\n\t\t\t<\/div>\n\t\t<\/a>\n\t\t<div class=\"expand-body js-expand-content\">\n\t\t\t<div class=\"expand-body-content\">\n\t\t\t\t<p>This study cycle promotes the development of competences in theoretical advancement and critical analysis, the acquisition of advanced research techniques both in the laboratory and in the field, and the production of original research work to be presented in the form of scientific articles or posters, oral presentations at scientific meetings, or meetings with decision-makers, managers, or science funders. MARE, rated Excellent in the latest evaluation by the Portuguese Foundation for Science and Technology (FCT), supports this study cycle by integrating its PhD researchers into the course units.<\/p>\n\t\t\t\t\n\t\t\t\t<!-- Plano de estudos -->\n\t\t\t\t\n\t\t\t<\/div>\n\t\t<\/div>\n\t\t\n\t\n\t\t<a href=\"#\" class=\"expand-header js-expand\">\n\t\t\t<div class=\"expand-header-text\">\n\t\t\t\t<h2 class=\"expand-block--title\">Course Duration<\/h2>\n\t\t\t<\/div>\n\t\t<\/a>\n\t\t<div class=\"expand-body js-expand-content\">\n\t\t\t<div class=\"expand-body-content\">\n\t\t\t\t<p>The Doctoral Study Cycle has a duration of 4 academic years (8 semesters), corresponding to a total of 240 ECTS credits.<\/p>\n\t\t\t\t\n\t\t\t\t<!-- Plano de estudos -->\n\t\t\t\t\n\t\t\t<\/div>\n\t\t<\/div>\n\t\t\n\t\n\t\t<a href=\"#\" class=\"expand-header js-expand\">\n\t\t\t<div class=\"expand-header-text\">\n\t\t\t\t<h2 class=\"expand-block--title\">Programme Objectives<\/h2>\n\t\t\t<\/div>\n\t\t<\/a>\n\t\t<div class=\"expand-body js-expand-content\">\n\t\t\t<div class=\"expand-body-content\">\n\t\t\t\t<ul>\n<li>To acquire an in-depth understanding and integration of the main theories of modern Biology and to apply professionally and competently the principal methods and techniques, particularly those relating to Behavioural Biology;<\/li>\n<li>To master advanced statistical analysis of ecological, evolutionary, behavioural, and genetic data, and the interpretation of results obtained;<\/li>\n<li>To design, plan, adapt, and conduct scientific research in Behavioural Biology, respecting the ethical principles underlying the research process and the use of animals in experimentation;<\/li>\n<li>To communicate research findings and their theoretical and methodological foundations clearly and professionally to diverse audiences, including scientists, members of the academic community, decision-makers, managers, and the public.<\/li>\n<\/ul>\n\t\t\t\t\n\t\t\t\t<!-- Plano de estudos -->\n\t\t\t\t\n\t\t\t<\/div>\n\t\t<\/div>\n\t\t\n\t\n\t\t<a href=\"#\" class=\"expand-header js-expand\">\n\t\t\t<div class=\"expand-header-text\">\n\t\t\t\t<h2 class=\"expand-block--title\">Admission<\/h2>\n\t\t\t<\/div>\n\t\t<\/a>\n\t\t<div class=\"expand-body js-expand-content\">\n\t\t\t<div class=\"expand-body-content\">\n\t\t\t\t<p>1 \u2013 The following may apply to Ispa\u2019s Doctoral Programmes:<\/p>\n<ol>\n<li>a) holders of a master\u2019s degree or legally equivalent qualification;<\/li>\n<li>b) holders of a bachelor\u2019s degree who possess an especially relevant academic or scientific curriculum recognised by Ispa\u2019s Scientific Council as attesting the ability to successfully complete this study cycle;<\/li>\n<li>c) holders of an academic, scientific, or professional curriculum recognised by Ispa\u2019s Scientific Council as attesting the ability to successfully complete this study cycle.<\/li>\n<\/ol>\n<p>2 \u2013 The recognition referred to in points b) and c) of paragraph 1 grants access only to the study cycle leading to the doctoral degree, and does not confer equivalence to, or recognition of, a bachelor\u2019s or master\u2019s degree.<\/p>\n<p>Please consult the regulation <a href=\"http:\/\/www.ispa.pt\/ficheiros\/documentos\/rg088_acesso_3ciclos.pdf\">here.<\/a><\/p>\n\t\t\t\t\n\t\t\t\t<!-- Plano de estudos -->\n\t\t\t\t\n\t\t\t<\/div>\n\t\t<\/div>\n\t\t\n\t\n\t\t<a href=\"#\" class=\"expand-header js-expand\">\n\t\t\t<div class=\"expand-header-text\">\n\t\t\t\t<h2 class=\"expand-block--title\">Timetable<\/h2>\n\t\t\t<\/div>\n\t\t<\/a>\n\t\t<div class=\"expand-body js-expand-content\">\n\t\t\t<div class=\"expand-body-content\">\n\t\t\t\t<p>Daytime.<\/p>\n\t\t\t\t\n\t\t\t\t<!-- Plano de estudos -->\n\t\t\t\t\n\t\t\t<\/div>\n\t\t<\/div>\n\t\t\n\t\n\t\t<a href=\"#\" class=\"expand-header js-expand\">\n\t\t\t<div class=\"expand-header-text\">\n\t\t\t\t<h2 class=\"expand-block--title\">Tuition Fees<\/h2>\n\t\t\t<\/div>\n\t\t<\/a>\n\t\t<div class=\"expand-body js-expand-content\">\n\t\t\t<div class=\"expand-body-content\">\n\t\t\t\t<p><strong><a href=\"https:\/\/intranet.ispa.pt\/inscricoes\/taxas-propinas\">Fees and tuition<\/a><br \/>\n<a href=\"https:\/\/intranet.ispa.pt\/content\/emolumentos-prazos-e-condi%C3%A7%C3%B5es-e-emiss%C3%A3o-de-documentos\">Charges and documents<\/a><br \/>\n<a href=\"https:\/\/intranet.ispa.pt\/sites\/default\/files\/users\/10008\/rg051_tesouraria_v11.pdf\">Please refer to the Treasury Regulations<\/a><\/strong><\/p>\n\t\t\t\t\n\t\t\t\t<!-- Plano de estudos -->\n\t\t\t\t\n\t\t\t<\/div>\n\t\t<\/div>\n\t\t\n\t\n\n\n\n\n\t<!-- SIDE CONTENT -->\n\t<div class=\"side-content block-margin\">\n\t\t<h2 class=\"side-content-title\">Accreditation<\/h2>\n\t\t<div class=\"side-content-text\">\n\t\t\t<p>Study cycle accredited by the Agency for Assessment and Accreditation of Higher Education (A3ES).<\/p>\n<p><a href=\"https:\/\/si.a3es.pt\/sia3es\/page?stage=ConsultaPublicaProcesso&amp;processID=24076#none\">Process (A3ES)<strong>\u00a0<\/strong>ACEF\/2021\/1201366<\/a><\/p>\n<p><a href=\"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/10\/plano-de-estudos-doutoramento-biologia-comportamento.pdf\" target=\"_blank\" rel=\"noopener\">Study Plan (Official Gazette, (D.R. 2nd series)<\/a><\/p>\n\t\t<\/div>\n\t<\/div><!-- \/ SIDE CONTENT -->\n\n\n\n\n\n\t<\/div><!-- \/column -->\n\t\t\t\t\n\t<!-- sidebar -->\n\t<div class=\"the-content-sidebar\">\n\n\n\n\n\n\t<!-- DOWNLOAD FILE -->\n\t<div class=\"download-file page-limit-right block-margin\">\n\t\t<div class=\"download-file-bg\"><\/div>\n\t\t<div class=\"download-file-content\">\n\t\t\t<h2 class=\"download-file-title\">SABER MAIS<\/h2>\n\t\t<\/div>\n\t\t<div class=\"download-file-button\">\n\t\t\t<a class=\"button-download\" href=\"\" target=\"_blank\">download<\/a>\n\t\t<\/div>\n\t<\/div><!-- \/ DOWNLOAD FILE -->\n\n","protected":false},"featured_media":5890,"template":"","ensino":[156],"psicologia":[],"biociencias":[158],"educacao":[],"class_list":["post-25820","oferta-formativa","type-oferta-formativa","status-publish","has-post-thumbnail","hentry","ensino-doctoral-degrees","biociencias-doctoral-degrees"],"acf":[],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v26.1.1 - https:\/\/yoast.com\/wordpress\/plugins\/seo\/ -->\n<title>Doctoral Degrees - Behavioural Biology - Ispa \u2013 Instituto Universit\u00e1rio<\/title>\n<meta name=\"description\" content=\"Conhe\u00e7a o Doutoramento em Biologia do Comportamento, suportado pelo MARE que promove t\u00e9cnicas avan\u00e7adas e trabalhos de investiga\u00e7\u00e3o originais.\" \/>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/\" \/>\n<meta property=\"og:locale\" content=\"en_US\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Doctoral Degrees - Behavioural Biology - Ispa \u2013 Instituto Universit\u00e1rio\" \/>\n<meta property=\"og:description\" content=\"Conhe\u00e7a o Doutoramento em Biologia do Comportamento, suportado pelo MARE que promove t\u00e9cnicas avan\u00e7adas e trabalhos de investiga\u00e7\u00e3o originais.\" \/>\n<meta property=\"og:url\" content=\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/\" \/>\n<meta property=\"og:site_name\" content=\"Ispa \u2013 Instituto Universit\u00e1rio\" \/>\n<meta property=\"article:modified_time\" content=\"2025-11-03T16:27:57+00:00\" \/>\n<meta property=\"og:image\" content=\"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/03\/header-doutoramento-biologia-comportamento-ispa.png\" \/>\n\t<meta property=\"og:image:width\" content=\"2046\" \/>\n\t<meta property=\"og:image:height\" content=\"982\" \/>\n\t<meta property=\"og:image:type\" content=\"image\/png\" \/>\n<meta name=\"twitter:card\" content=\"summary_large_image\" \/>\n<meta name=\"twitter:label1\" content=\"Est. reading time\" \/>\n\t<meta name=\"twitter:data1\" content=\"1 minute\" \/>\n<script type=\"application\/ld+json\" class=\"yoast-schema-graph\">{\"@context\":\"https:\/\/schema.org\",\"@graph\":[{\"@type\":\"WebPage\",\"@id\":\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/\",\"url\":\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/\",\"name\":\"Doctoral Degrees - Behavioural Biology - Ispa \u2013 Instituto Universit\u00e1rio\",\"isPartOf\":{\"@id\":\"https:\/\/www.ispa.pt\/en\/#website\"},\"primaryImageOfPage\":{\"@id\":\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#primaryimage\"},\"image\":{\"@id\":\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#primaryimage\"},\"thumbnailUrl\":\"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/03\/header-doutoramento-biologia-comportamento-ispa.png\",\"datePublished\":\"2021-03-03T15:30:59+00:00\",\"dateModified\":\"2025-11-03T16:27:57+00:00\",\"description\":\"Conhe\u00e7a o Doutoramento em Biologia do Comportamento, suportado pelo MARE que promove t\u00e9cnicas avan\u00e7adas e trabalhos de investiga\u00e7\u00e3o originais.\",\"breadcrumb\":{\"@id\":\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#breadcrumb\"},\"inLanguage\":\"en-US\",\"potentialAction\":[{\"@type\":\"ReadAction\",\"target\":[\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/\"]}]},{\"@type\":\"ImageObject\",\"inLanguage\":\"en-US\",\"@id\":\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#primaryimage\",\"url\":\"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/03\/header-doutoramento-biologia-comportamento-ispa.png\",\"contentUrl\":\"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/03\/header-doutoramento-biologia-comportamento-ispa.png\",\"width\":2046,\"height\":982},{\"@type\":\"BreadcrumbList\",\"@id\":\"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#breadcrumb\",\"itemListElement\":[{\"@type\":\"ListItem\",\"position\":1,\"name\":\"Home\",\"item\":\"https:\/\/www.ispa.pt\/en\/\"},{\"@type\":\"ListItem\",\"position\":2,\"name\":\"Oferta formativa\",\"item\":\"https:\/\/www.ispa.pt\/en\/ensino\/academic-offer\/\"},{\"@type\":\"ListItem\",\"position\":3,\"name\":\"Behavioural Biology\"}]},{\"@type\":\"WebSite\",\"@id\":\"https:\/\/www.ispa.pt\/en\/#website\",\"url\":\"https:\/\/www.ispa.pt\/en\/\",\"name\":\"Ispa \u2013 Instituto Universit\u00e1rio\",\"description\":\"Institui\u00e7\u00e3o de refer\u00eancia nas \u00e1reas da Psicologia, Bioci\u00eancia e Educa\u00e7\u00e3o.\",\"publisher\":{\"@id\":\"https:\/\/www.ispa.pt\/en\/#organization\"},\"potentialAction\":[{\"@type\":\"SearchAction\",\"target\":{\"@type\":\"EntryPoint\",\"urlTemplate\":\"https:\/\/www.ispa.pt\/en\/?s={search_term_string}\"},\"query-input\":{\"@type\":\"PropertyValueSpecification\",\"valueRequired\":true,\"valueName\":\"search_term_string\"}}],\"inLanguage\":\"en-US\"},{\"@type\":\"Organization\",\"@id\":\"https:\/\/www.ispa.pt\/en\/#organization\",\"name\":\"Ispa \u2013 Instituto Universit\u00e1rio\",\"url\":\"https:\/\/www.ispa.pt\/en\/\",\"logo\":{\"@type\":\"ImageObject\",\"inLanguage\":\"en-US\",\"@id\":\"https:\/\/www.ispa.pt\/en\/#\/schema\/logo\/image\/\",\"url\":\"https:\/\/www.ispa.pt\/wp-content\/uploads\/2020\/03\/logo.svg\",\"contentUrl\":\"https:\/\/www.ispa.pt\/wp-content\/uploads\/2020\/03\/logo.svg\",\"width\":206,\"height\":72,\"caption\":\"Ispa \u2013 Instituto Universit\u00e1rio\"},\"image\":{\"@id\":\"https:\/\/www.ispa.pt\/en\/#\/schema\/logo\/image\/\"}}]}<\/script>\n<!-- \/ Yoast SEO plugin. -->","yoast_head_json":{"title":"Doctoral Degrees - Behavioural Biology - Ispa \u2013 Instituto Universit\u00e1rio","description":"Conhe\u00e7a o Doutoramento em Biologia do Comportamento, suportado pelo MARE que promove t\u00e9cnicas avan\u00e7adas e trabalhos de investiga\u00e7\u00e3o originais.","robots":{"index":"index","follow":"follow","max-snippet":"max-snippet:-1","max-image-preview":"max-image-preview:large","max-video-preview":"max-video-preview:-1"},"canonical":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/","og_locale":"en_US","og_type":"article","og_title":"Doctoral Degrees - Behavioural Biology - Ispa \u2013 Instituto Universit\u00e1rio","og_description":"Conhe\u00e7a o Doutoramento em Biologia do Comportamento, suportado pelo MARE que promove t\u00e9cnicas avan\u00e7adas e trabalhos de investiga\u00e7\u00e3o originais.","og_url":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/","og_site_name":"Ispa \u2013 Instituto Universit\u00e1rio","article_modified_time":"2025-11-03T16:27:57+00:00","og_image":[{"width":2046,"height":982,"url":"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/03\/header-doutoramento-biologia-comportamento-ispa.png","type":"image\/png"}],"twitter_card":"summary_large_image","twitter_misc":{"Est. reading time":"1 minute"},"schema":{"@context":"https:\/\/schema.org","@graph":[{"@type":"WebPage","@id":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/","url":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/","name":"Doctoral Degrees - Behavioural Biology - Ispa \u2013 Instituto Universit\u00e1rio","isPartOf":{"@id":"https:\/\/www.ispa.pt\/en\/#website"},"primaryImageOfPage":{"@id":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#primaryimage"},"image":{"@id":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#primaryimage"},"thumbnailUrl":"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/03\/header-doutoramento-biologia-comportamento-ispa.png","datePublished":"2021-03-03T15:30:59+00:00","dateModified":"2025-11-03T16:27:57+00:00","description":"Conhe\u00e7a o Doutoramento em Biologia do Comportamento, suportado pelo MARE que promove t\u00e9cnicas avan\u00e7adas e trabalhos de investiga\u00e7\u00e3o originais.","breadcrumb":{"@id":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#breadcrumb"},"inLanguage":"en-US","potentialAction":[{"@type":"ReadAction","target":["https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/"]}]},{"@type":"ImageObject","inLanguage":"en-US","@id":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#primaryimage","url":"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/03\/header-doutoramento-biologia-comportamento-ispa.png","contentUrl":"https:\/\/www.ispa.pt\/wp-content\/uploads\/2021\/03\/header-doutoramento-biologia-comportamento-ispa.png","width":2046,"height":982},{"@type":"BreadcrumbList","@id":"https:\/\/www.ispa.pt\/en\/academic-offer\/doctors-behavioural-biology\/#breadcrumb","itemListElement":[{"@type":"ListItem","position":1,"name":"Home","item":"https:\/\/www.ispa.pt\/en\/"},{"@type":"ListItem","position":2,"name":"Oferta formativa","item":"https:\/\/www.ispa.pt\/en\/ensino\/academic-offer\/"},{"@type":"ListItem","position":3,"name":"Behavioural Biology"}]},{"@type":"WebSite","@id":"https:\/\/www.ispa.pt\/en\/#website","url":"https:\/\/www.ispa.pt\/en\/","name":"Ispa \u2013 Instituto Universit\u00e1rio","description":"Institui\u00e7\u00e3o de refer\u00eancia nas \u00e1reas da Psicologia, Bioci\u00eancia e Educa\u00e7\u00e3o.","publisher":{"@id":"https:\/\/www.ispa.pt\/en\/#organization"},"potentialAction":[{"@type":"SearchAction","target":{"@type":"EntryPoint","urlTemplate":"https:\/\/www.ispa.pt\/en\/?s={search_term_string}"},"query-input":{"@type":"PropertyValueSpecification","valueRequired":true,"valueName":"search_term_string"}}],"inLanguage":"en-US"},{"@type":"Organization","@id":"https:\/\/www.ispa.pt\/en\/#organization","name":"Ispa \u2013 Instituto Universit\u00e1rio","url":"https:\/\/www.ispa.pt\/en\/","logo":{"@type":"ImageObject","inLanguage":"en-US","@id":"https:\/\/www.ispa.pt\/en\/#\/schema\/logo\/image\/","url":"https:\/\/www.ispa.pt\/wp-content\/uploads\/2020\/03\/logo.svg","contentUrl":"https:\/\/www.ispa.pt\/wp-content\/uploads\/2020\/03\/logo.svg","width":206,"height":72,"caption":"Ispa \u2013 Instituto Universit\u00e1rio"},"image":{"@id":"https:\/\/www.ispa.pt\/en\/#\/schema\/logo\/image\/"}}]}},"_links":{"self":[{"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/oferta-formativa\/25820","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/oferta-formativa"}],"about":[{"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/types\/oferta-formativa"}],"wp:featuredmedia":[{"embeddable":true,"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/media\/5890"}],"wp:attachment":[{"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/media?parent=25820"}],"wp:term":[{"taxonomy":"ensino","embeddable":true,"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/ensino?post=25820"},{"taxonomy":"psicologia","embeddable":true,"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/psicologia?post=25820"},{"taxonomy":"biociencias","embeddable":true,"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/biociencias?post=25820"},{"taxonomy":"educacao","embeddable":true,"href":"https:\/\/www.ispa.pt\/en\/wp-json\/wp\/v2\/educacao?post=25820"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}